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David Penney
University of Manchester
Summary
I have had a life-long fascination with all things creepy crawly and still vividly remember exactly how this came about. My qualifications include a BSc in Zoology and a PhD in amber palaeobiology and I am a Fellow of the Royal Entomological Society. Following my PhD I worked as Assistant Keeper of Zoology at Manchester Museum for 18 months, before embarking on a post-doc at the University of Manchester studying fossil spiders. Following completion of the post-doc I have retained honorary research status at the University of Manchester, most recently as an Honorary Lecturer in the Faculty of Life Sciences. I have extensive experience of ecology and biodiversity fieldwork in many different countries, including the tropics and taught on residential ecology field courses for 15 years. I also run the scientific publishing company Siri Scientific Press, part of Siri Scientific Services, which also offers consultancy services in a wide range of areas including fossil and living spider and insect identification (please email me with any book or other queries).
Research interests
My research interests focus on extant and fossil spiders and insects. They cover a broad spectrum taxonomically, geographically, geologically and theoretically. They include: extant and fossil spider taxonomy and systematics; evolutionary history of spiders; the qualitative and quantitative use of data in testing hypotheses of palaeoecology and palaeogeography; preservation bias of organisms preserved in amber; reconstruction of fossil ecosystems; the effects of mass extinctions on terrestrial invertebrate faunas; the timing of the radiations of the major extant spider families; the completeness versus the adequacy of the spider fossil record; predator–prey co-evolutionary processes; and the origins and current biodiversity of the Greater Antillean spider fauna. I am particularly interested in the application of amber derived data in quantitative palaeobiological studies of broad interest. Some highlights include demonstrating that amber is biased to preserving active, trunk-dwelling faunas (Paleobiology 2002), that spiders survived the end-Cretaceous mass extinction event (Evolution 2003), that spiders co-radiated alongside their insect prey over geological time (Trans. R. Soc. Edinburgh, Earth Sci. 2004), that extant and fossil Hispaniolan spider assemblages are directly comparable (Palaeo3 2005) and have South American origins (Book: Dominican Amber Spiders, 2008), that different amber faunas retain ecological information and are directly comparable in this respect (Royal Society, Biology Letters 2006), description of the oldest orb-weaving spider (Royal Society, Biology Letters 2006), the novel application of VHR computed tomography to amber inclusions (Zootaxa 2007, 2011; Naturwissenschaften 2011; Royal Society, Biology Letters 2011), a full review of the amber and non-amber spider fossil record (Biological Reviews 2010 – updated and illustrated in my 2011 monograph book on Fossil Spiders; with summary statistics of palaeospecies taxonomy ZooKeys 2012), erection of a new fossil spider family (Bull. Brit. Arachnol. Soc. 2011), and a comprehensive review of insect behaviour in the fossil record (Biological Reviews – submitted). I co-author (with Jason Dunlop in Berlin) the World Fossil Arachnid Species Checklist which is hosted on the World Spider Catalog website and is updated every six months, and we recently (2012) co-authored a full colour monograph book on Fossil Arachnids. I am also interested in extant faunas, where my recent research has focused on the biodiversity and ecology of arachnids and insects in The Gambia, West Africa, resulting in the publication of several field guides and scientific papers, and also on epiphytic invertebrate communities in the Neotropics (Philosophical Transactions of the Royal Society 2011).
I am currently working on the descriptions of various new fossil taxa (insects and arachnids) in both amber and non-amber deposits, a remarkable case of phoresy in the fossil record, a phenological and ecological study of extant Nephila spiders, and various X-ray computed tomography studies, in addition to editing a multi-authored volume on spider research in the 21st century and writing a book on fossil insects, with various others also under consideration. I am currently undertaking a short (May-July 2012) project (with Prof. Terry Brown & Dr Richard Preziosi) to attempt DNA extraction from subfossilized bees Trigonisca sp. (Apidae: Apinae: Meliponini) in Colombian copal using next generation sequencing techniques. We will also include a Dominican amber sample in the analysis just on the off chance! Although this 2-month DNA project is funded, the rest of my research is not. If you would like to sponsor my research and have a new species named after you for doing so please drop me an email.
| Current Institution | University of Manchester |
| Current School | Faculty of Life Sciences |
| Department | I also run Siri Scientific Press: http://www.siriscientificpress.co.uk |
| Disciplines | |
| Address | Manchester Lancashire United Kingdom Phone: |
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Education
Work Experience
Achievements
General
Publication Summary
1. Authored books
- B10. Penney, D. & Jepson, J.E. (in prep. for 2013) Fossil Insects. Siri Scientific Press, Manchester.
- B9. Penney, D. (2012, in press) Field Guide to Wildlife of The Gambia 2nd edition, 160 pp. Siri Scientific Press, Manchester.
- B8. Dunlop, J.A. & Penney, D. (2012) Fossil Arachnids, 192 pp. Monograph Series, Volume 2. Siri Scientific Press, Manchester.
- B7. Penney, D. & Green, D. (2011) Fossils in Amber: snapshots of prehistoric forest life, 226 pp. Siri Scientific Press, Manchester.
- B6. Penney, D. & Selden, P.A. (2011) Fossil Spiders: the evolutionary history of a mega-diverse order, 128 pp. Monograph Series, Volume 1. Siri Scientific Press, Manchester.
- B5. Penney, D. (2010) Amber, 28 pp. Siri Scientific Press, Manchester.
- B4. Penney, D. (2009) Field Guide to Wildlife of The Gambia, 120 pp. Siri Scientific Press, Manchester.
- B3. Penney, D. (2009) Common Spiders and Other Arachnids of The Gambia, West Africa, 80 pp. Siri Scientific Press, Manchester.
- B2. Penney, D. (2009) Field Guide to Butterflies of The Gambia, West Africa, 80 pp. Siri Scientific Press, Manchester.
- B1. Penney, D. (2008) Dominican Amber Spiders: a comparative palaeontological-neontological approach to identification, faunistics, ecology and biogeography, 178 pp. Siri Scientific Press, Manchester.
2. Edited volumes
- E4. Penney, D. (ed.) (in prep. for 2012/13) Spider research in the 21st century. Siri Scientific Press, Manchester.
- E3. Coram, R. & Jepson, J.E. (Penney, D., ed.) (2012) Fossil insects of the Purbeck Formation of Southern England: palaeoentomology from the dawn of the Cretaceous, 144 pp. Monograph Series, Volume 3. Siri Scientific Press, Manchester.
- E2. Penney, D. (ed.) (2010) Biodiversity of fossils in amber from the major world deposits, 304 pp. Siri Scientific Press, Manchester.
- E1. Logunov, D.V. & Penney, D. (eds) (2004) European Arachnology 2003. Proceedings of the 21st European Colloquium of Arachnology (St. Petersburg, Russia, August 2003). KMK Scientific Press, Moscow, 374 pp.
3. Chapters in books
- C7. Penney, D. (in press) Palaeontology. In Penney, D. (ed.) Spider research in the 21st century. Siri Scientific Press, Manchester.
- C6. Dunlop, J.A., Penney, D. & Jekel, D. (2012) A summary list of fossil spiders and their relatives. 261 pp. In:Platnick, N.I. 2012. The world spider catalog, version 12.5. American Museum of Natural History, online at http://research.amnh.org/entomology/spiders/catalog/index.html
- C5. Dunlop, J.A., Penney, D. & Jekel, D. (2011) A summary list of fossil spiders and their relatives. 253 pp. In:Platnick, N.I. 2011. The world spider catalog, version 12.0. American Museum of Natural History, online at http://research.amnh.org/entomology/spiders/catalog/index.html
- C4. Penney, D. (2010) Dominican amber.Pp. 22–41 In: Penney, D. (ed.) Biodiversity of fossils in amber from the major world deposits.Siri Scientific Press, Manchester.
- C3. Penney, D. & Green, D. (2010) Introduction, preparation, study and conservation of amber inclusions. Pp. 5–21 In:Penney, D. (ed.) Biodiversity of fossils in amber from the major world deposits. Siri Scientific Press, Manchester.
- C2. Penney, D. & Preziosi, R.F. (2010) On inclusions in subfossil resins (copal). Pp. 299–303 In:Penney, D. (ed.) Biodiversity of fossils in amber from the major world deposits. Siri Scientific Press, Manchester.
- C1. Penney, D. (2007) Hispaniolan spider biodiversity and the importance of combining neontological and palaeontological data in analyses of historical biogeography. Pp. 63–100 In: Schwartz, J. (ed.) Focus on Biodiversity Research. Nova Science Publishers Inc., New York. Invited submission.
4. Academic journal papers (reverse chronological order) pdfs available on request
- R74. Penney, D. & Ostapenko, K. (in review) The fossil record of insect behaviour. Biological Reviews.
- R73. Penney, D., McNeil, A., Green, D.I., Bradley, R., Jepson, J.E., Whithers, P.J. & Preziosi, R.F. (in prep.) Unique Collembola–Ephemeroptera phoretic behaviour fossilized in amber. For Nature.
- R72. Penney, D., Green, D.I., McNeil, A., Bradley, R., Marusik, Y., Whithers, P.J. & Preziosi, R.F. (in press) A new species of Craspedisia (Araneae: Theridiidae) in Miocene Dominican amber, imaged using X-ray computed tomography. Paleontological Journal.
- R71. Penney, D., Dunlop, J.A. & Marusik, Y.M. (2012) Summary statistics for fossil spider species taxonomy. ZooKeys, 192: 1–13.
- R70. Penney, D. (in press) Biodiversity of fossils in amber from the major world deposits. Mitteilungen DgaEE.
- R69. Penney, D., Green, D.I., Titchener, S.B, Titchener, B.G., Brown, T.A. & Preziosi, R.F. (2012) An unusual palaeobiocoenosis of (sub)fossil spiders in Colombian copal. Bulletin of the British Arachnological Society, 15: 241–244.
- R68. Dunlop, J.A. & Penney, D. (2011) Order Araneae Clerck, 1757. In: Zhang, Z.-Q. (ed.) Animal biodiversity: An outline of higher-level classification and survey of taxonomic richness. Zootaxa, 3148, 149–153.
- R67. Dunlop, J.A., Wirth, S., Penney, D., McNeil, A., Bradley, R.S.,Withers, P.J. & Preziosi, R.F. (2011) A minute fossil phoretic mite recovered by X-ray computed tomography. Biology Letters, doi 10.1098/rspb.2011.0923.
- R66. Jepson, J., Penney, D. & Green, D.I. (2011) First fossil Phatnoma (Heteroptera: Tingidae: Cantacaderinae), in Miocene amber from the Dominican Republic. Zootaxa, 2975, 59–63.
- R65. Dunlop, J.A., Penney, D., Daluge, N., Jager, P., McNeil, A., Bradley, R., Whithers, P.J. & Preziosi, R.F. (2011) Computed tomography recovers data from historical amber: an example from huntsman spiders. Naturwissenschaften, DOI 10.1007/s00114-011-0796-x. Selected as cover article.
- R64. Penney, D. (2011) Grandoculidae: a new fossil spider family from the Upper Cretaceous of Canada. Bulletin of the British Arachnological Society,15, 179–180.
- R63. Penney, D., Green, D.I., McNeil, A., Bradley, R., Marusik, Y.M., Withers, P.J. & Preziosi, R.F. (2011) A new species of anapid spider (Arthropoda: Araneae, Anapidae) in Eocene Baltic amber, imaged using X-ray computed tomography. Zootaxa, 2742, 61–68.
- R62. Zytynska, S.E., Fay, M.F., Penney, D. & Preziosi, R.F. (2011) Genetic variation in a tropical tree species influences the structure of associated epiphytic plant and invertebrate communities in a complex forest ecosystem. Philosophical Transactions of the Royal Society B., 366, 1329–1336.
- R61. McNeil, A., Bradley, R.S., Withers, P.J. & Penney, D. (2010) Imaging Fossilised Spiders in Amber Using Lab-Based Phase Contrast X-ray Tomography.In:Stock, S.R (ed.) Developments in X-Ray Tomography VII. Proc. SPIE, San Diego, 7804, 78041Q1–7.
- R60. Penney, D. (2010) The evolution of jumping spiders (Araneae: Salticidae): the palaeontological evidence. Peckhamia, 81.8, 1–3.
- R59. Penney, D. & Langan, A.M. (2010) Morphometric identification of fossils spiders: comment. Paleontological Journal, 44, 644–648. Original Russian text published in Paleontologicheskii Zhurnal, 44, 44–48.
- R58. Jepson, J., Penney, D. & Green, D.I. (2010) A new species of brown lacewing from Eocene Baltic amber (Insecta: Neuroptera). Zootaxa, 2692, 61–68.
- R57. Saupe, E.E., Selden, P.A. & Penney, D. (2010) First fossil Molinaranea (Araneae: Araneidae), from Middle Miocene Dominican amber, with a phylogenic and paleobiogeographic analysis of the genus. Zoological Journal of the Linnean Society, 158, 711–725.
- R56. Selden, P.A. & Penney, D. (2010) Fossil spiders. Biological Reviews, 85, 171–206. Invited paper. Selected as cover article.
- R55. Penney, D. (2009) A new spider family record for Hispaniola—a new species of Plectreurys (Araneae: Plectreuridae) in Miocene Dominican amber. Zootaxa, 2144, 65–68.
- R54. Penney, D. & Gabriel, R.A. (2009) Feeding behavior of trunk-living jumping spiders (Salticidae) in a coastal primary forest in The Gambia. Journal of Arachnology, 37, 113–115.
- R53. Penney, D., Marusik, Y., Wheater, C.P. & Langan, A.M. (2009) First Gambian Ricinulei and the northernmost African record for the order (Arachnida: Ricinulei: Ricinioididae). Zootaxa, 2021, 66–68.
- R52. Dunlop, J.A. & Penney, D. (2009) Case 3475 Myrmarachne MacLeay, 1839 (Araneae, Salticidae): proposed conservation of the generic name. Bulletin of Zoological Nomenclature, 66, 20–23.
- R51. Selden, P.A. & Penney, D. (2009) A fossil spider (Araneae: Pisauridae) of Eocene age from Horsefly, British Columbia, Canada. Contributions to Natural History, 12, 1269–1282.
- R50. Penney, D. (2008) Possible anti-predator function for the stabilimentum in a West African (Bijilo Forest, The Gambia) Cyclosa (Araneae, Araneidae). Journal of Afrotropical Zoology, 4, 143–146.
- R49. Dunlop, J.A.,Penney, D., Tetlie, O.E. & Anderson, L.I. (2008) How many species of fossil arachnid are there? Journal of Arachnology, 36, 267–272. Invited paper for 17th International Congress of Arachnology, Brazil (2007)
- R48. Dunlop, J.A., Harms, D. &Penney, D. (2008) A fossil tarantula (Araneae: Theraphosidae) from Miocene Chiapas amber, Mexico. Revista Iberica de Aracnologia,15, 9–17.
- R47. Langan, A.M., Shuker, D.M., Cullen, W.R.,Penney, D., Preziosi, R.F. & Wheater, C.P. 2008. Relationships between student characteristics and self-, peer and tutor evaluations of oral presentations. Assessment and Evaluation in Higher Education, 33, 179–193.
- R46. Penney, D. (2007) A new fossil oonopid spider, in lowermost Eocene amber from the Paris Basin, with comments on the fossil spider assemblage. African
- Invertebrates, 48, 71–75.
- R45. Penney, D. (2007 for 2006) The oldest fossil pholcid and selenopid spiders, in lowermost Eocene amber from the Paris Basin, France. Journal of Arachnology, 34, 592–598.
- R44. Penney, D. & Selden, P.A. (2007) Fossils explained: Spinning with the dinosaurs: the fossil record of spiders. Geology Today 23, 231–237. Invited paper
- R43. Penney, D., Dierick, M., Cnudde, V., Masschaele, B., Vlasssenbroeck, J., Van Hoorebeke, L. & Jacobs, P. (2007) First fossil Micropholcommatidae (Araneae), imaged in Eocene Paris amber using X-Ray Computed Tomography. Zootaxa, 1623, 47–53.
- R42. Jepson, J.E. & Penney, D. (2007) Neuropteran (Insecta) palaeodiversity with predictions for the Cretaceous fauna of the Wealden. Palaeogeography Palaeoclimatology Palaeoecology, 248, 109–118.
- R41. Bennett, S.P. & Penney, D. (2006) New spider (Araneae) species records for the Isle of Mull, UK. Acta Zoologica Bulgarica, supplement 1, 3pp.
- R40. Penney, D. (2006a for 2005) An annotated systematic catalogue, including synonymies and transfers, of Miocene Dominican Republic amber spiders described up until 2005.Revista Iberica de Aracnologia 12, 25–52.
- R39. Penney, D. (2006b) Fossil oonopid spiders in Cretaceous ambers from Canada and Myanmar. Palaeontology, 49, 229–235.
- R38. Penney, D. (2006c) Tertiary Neotropical Hersiliidae (Arthropoda, Araneae) with new combinations for the extant fauna and comments on historical biogeography of the family. Palaeontology, 49, 899–906.
- R37. Penney, D. (2006d) The oldest lagonomegopid spider, a new species in Lower Cretaceous amber from Álava, Spain. Geologica Acta, 4, 377–382.
- R36. Penney, D. (2006e) Fossils in amber: unlocking the secrets of the past. Biologist, 53, 247–251. Invited paper
- R35. Penney, D. & Bennet, S.P. (2006) First unequivocal mermithid–linyphiid parasite–host association. Journal of Arachnology, 34, 273–278.
- R34. Penney, D. & Langan, A.M. (2006) Comparing amber fossils across the Cenozoic. Biology Letters, 2, 266–270.doi:10.1098/rsbl.2006.0442
- R33. Penney, D. & Ortuño, V.M. (2006) Oldest true orb-weaving spider (Araneae: Araneidae). Biology Letters, 2, 447–450.
- R32. Penney, D. & Selden, P.A. (2006) First fossil Huttoniidae (Araneae), in Late Cretaceous Canadian Cedar and Grassy Lake ambers. Cretaceous Research, 27, 442–446.
- R31. Penney, D. & Selden, P.A. (2006) Assembling the Tree of Life—Phylogeny of Spiders: a review of the strictly fossil spider families. Acta Zoologica Bulgarica, supplement 1, 25–39.
- R30. Penney, D. (2005a) First fossil Filistatidae: A new species of Misionella in Miocene amber from the Dominican Republic. Journal of Arachnology, 33, 93–100.
- R29. Penney, D. (2005b) First Caribbean Floricomus(Araneae: Linyphiidae), a new species in Miocene Dominican Republic amber. A new synonymy for the extant American fauna. Geologica Acta, 3, 59–64.
- R28. Penney, D.(2005c) Fossil blood droplets in Miocene Dominican amber yield clues to speed and direction of resin secretion. Palaeontology, 48, 925–927.
- R27. Penney, D. (2005d) Importance of Dominican Republic Amber for determining taxonomic bias of fossil resin preservation—A case study of spiders. Palaeogeography Palaeoclimatology Palaeoecology, 223, 1–8.
- R26. Penney, D. (2005e) The fossil spider family Lagonomegopidae in Cretaceous ambers with description of a new genus and species from Myanmar. Journal of Arachnology, 33, 439–444.
- R25. Langan, A.M., Wheater, C.P., Preziosi, R.F., Boyle, J., Haines, B.J., Shaw, E.M., Penney, D., Cullen, W.R., Oldekopf, J., Aschcroft, C. & Lockey, L. (2005) Peer assessment of student presentations: effects of gender, cross-University affiliation and participation in the development of assessment criteria. Assessment and Evaluation in Higher Education, 30, 19–32.
- R24.Penney, D., Ono, H. & Selden, P.A. (2005) A new synonymy for the Madagascan copal spider fauna (Araneae, Selenopidae). Journal of Afrotropical Zoology, 2, 43–46.
- R23. Penney, D. (2004a) A new genus and species of Pisauridae (Araneae) in Cretaceous Burmese amber. Journal of Systematic Palaeontology, 2, 141–145, pl. 4. Invited paper. New journal for 2003, formerly Bull. Nat. Hist. Mus. London, Geol. Ser.
- R22. Penney, D. (2004b) New extant and fossil Dominican Republic spider records, with two new synonymies and comments on taphonomic bias of amber preservation. Revista Iberica de Aracnologia, 9, 183–190.
- R21. Penney, D. (2004c) New spiders in Upper Cretaceous amber from New Jersey in the American Museum of Natural History (Arthropoda, Araneae). Palaeontology, 47, 367–375.
- R20. Penney, D. (2004d) Does the fossil record of spiders track that of their principal prey, the insects? Transactions of the Royal Society of Edinburgh, Earth Sciences, 94, 275–281. Invited paper
- R19. Penney, D. (2004e) Cretaceous Canadian amber spider and the palpimanoidean nature of lagonomegopids. Acta Palaeontologica Polonica, 49, 579–584.
- R18. Penney, D. (2004f) New species records of Peucetia (Oxyopidae) from Uganda, with comments on the taxonomy of some African species. Bulletin of the British Arachnological Society, 13, 93–94.
- R17. Penney, D. (2004g) Estimations of global extant spider family richness using neontological and palaeontological data. In Logunov, D.V. & Penney, D. (eds). European Arachnology 2003. Proceedings of the 21st European Colloquium of Arachnology (St. Petersburg, Russia, August 2003). KMK Scientific Press, Moscow, 225–234.
- R16. Marusik, Y.M. & Penney, D. (2004) A survey of Baltic amber Theridiidae (Araneae) inclusions with descriptions of six new species. In Logunov, D.V. & Penney, D. (eds). European Arachnology 2003. Proceedings of the 21st European Colloquium of Arachnology (St. Petersburg, Russia, August 2003). KMK Scientific Press, Moscow, 201–218. Invited paper
- R15. Platnick, N.I. & Penney, D. (2004) A revision of the widespread spider genusZimiris (Araneae, Prodidomidae). American Museum Novitates, 3450, 1–12.
- R14. Penney, D. (2003a) A new deinopoid spider from Cretaceous Lebanese amber. Acta Palaeontologica Polonica, 48, 569–574.
- R13. Penney, D.(2003b) Afrarchaea grimaldii, a new species of Archaeidae (Araneae) in Cretaceous Burmese amber. Journal of Arachnology, 31, 122–130.
- R12. Penney, D., Wheater, C.P. & Selden, P.A. (2003) Resistance of spiders to Cretaceous–Tertiary extinction events. Evolution, 57, 2599–2607.
- R11. García-Villafuerte, M.A. & Penney, D. (2003) Lyssomanes Hentz (Araneae, Salticidae) in Oligocene–Miocene Chiapas amber. Journal of Arachnology, 31, 400–404.
- R10. Menon, F., Penney, D., Selden, P.A. & Martill, D.M. (2003) A new fossil scolopendromorph centipede from the Crato Formation of Brazil. Bulletin of the British Myriapod and Isopod Group, 9, 62–66.
- R9. Selden, P.A. & Penney, D. (2003) Lower Cretaceous spiders (Arthropoda, Arachnida, Araneae) from Spain. Neues Jahrbuch für Geologie und Paläontologie Montashefte Stuttgart, 2003,175–192.
- R8. Penney, D. (2002a) Spiders in Upper Cretaceous amber from New Jersey (Arthropoda, Araneae). Palaeontology, 45, 709–724.
- R7. Penney, D. (2002b) Paleoecology of Dominican amber preservation—spider (Araneae) inclusions demonstrate a bias for active, trunk-dwelling faunas. Paleobiology, 28, 389–398.
- R6. Penney, D. & Perez-Gelabert, D.E. (2002) Comparison of the Recent and Miocene Hispaniolan spider faunas. Revista Iberica de Aracnologia, 6, 203–223.
- R5. Penney, D. & Selden, P.A. (2002) The oldest linyphiid spider, in Lower Cretaceous Lebanese amber (Araneae, Linyphiidae, Linyphiinae). Journal of Arachnology, 30, 487–493.
- R4. Penney, D. (2001) Advances in the taxonomy of spiders in Miocene amber from the Dominican Republic (Arthropoda, Araneae). Palaeontology, 44, 987–1009.
- R3. Penney, D. (2000a) Miocene spiders in Dominican amber (Oonopidae, Mysmenidae). Palaeontology, 43, 343–357.
- R2. Penney, D. (2000b) Anyphaenidae in Miocene Dominican Republic amber (Arthropoda: Araneae). Journal of Arachnology, 28, 223–226.
- R1. Penney, D. (1999) Hypotheses for the Recent Hispaniolan spider fauna based on the Dominican Republic amber spider fauna. Journal of Arachnology, 27, 64–70.
5. Conference abstracts
- A40. Marusik, Y.M. & Penney, D. (2012) Conformation of the male palp in Dictynidae (Araneae) and related taxa. Abstracts. 14th Russian Entolomological Congress, St. Petersburg, in press.
- A39. Marusik, Y.M. & Penney, D. (2012) Terminology of the copulatory organs in spiders (Arachnida: Araneae): chaos or order? Abstracts. 14th Russian Entolomological Congress, St. Petersburg, in press.
- A38. Penney, D., Dunlop, J.A. & Marusik, Y.M. (2012) Summary statistics of fossil spider species taxonomy. Abstracts. 27th European Congress of Arachnology, Slovenia.
- A37. Penney, D. (2011) Spider palaeontology in the 21st century. British Arachnological Society AGM, unpublished. Invited speaker.
- A36. Dunlop, J.A., Garwood, R. & Penney, D. (2011) The devil in the detail: modern morphological methods allow the confident placement of fossil arachnids as (molecular) calibration points. Abstracts. 8th Biennial Meeting of the Systematics Association, Belfast, 43. Invited lecture.
- A35. Dunlop, J.A., Penney, D., Daluge, N., Jager, P., McNeil, A., Bradley, R. & Whithers, P.J. (2011) Computed tomography recovers data from historical amber: an example from huntsman spiders. Entomologentag de Deutsche Entomologische Gesellschaft, Humboldt University, Berlin, 223.
- A34. Penney, D. (2011) Biodiversity of fossils in amber from the major world deposits. Entomologentag de Deutsche Entomologische Gesellschaft, Humboldt University, Berlin, 214. Invited keynote. Abstract in DgaaE Nachrichten, 25: 120.
- A33. Dunlop, J.A. & Penney, D. (2010) The arachnid fossil record: progress and prospects.Abstracts. PalAss AM, Ghent, Belgium, 20.
- A32. Penney, D., Marusik, Y.M., Green, D.I., McNeil, A., Bradley, R., Withers, P.J. & Preziosi, R.F. (2010) X-ray computed tomography of a new Craspedisia (Araneae: Theridiidae) in Miocene Dominican amber. Extended Abstracts. XVIII Int. Cong. Arachnol., Poland, 331–334.
- A31. Marusik, Y.M. & Penney, D. (2010) Conformation of the male palp in some spiders belonging to the RTA-clade and problems in taxonomy. Abstracts. XVIII Int. Cong. Arachnol., Poland, 272–276.
- A30. McNeil, A. Bradley, R.S., Withers, P.J. & Penney, D. (2010) Imaging fossilised spiders in amber using lab-based phase contrast X-ray tomography. SPIE Optical Engineering + Applications, San Diego, California, Paper 7804-62 of Conference 7804. Abstracts online at: http://spie.org//app/program/index.cfm?event_id=894261&export_id=x13102&ID=x13188&redir=x13188.xml&search_text=amber&type=2&programDays=0&x=21&y=3
- A29. Saupe, E.E., Selden, P.A. & Penney, D. (2010) Using Dominican amber fossils to elucidate biogeographic patterns in the west indies: a case study with the genus Molinaranea (Araneae, Araneidae). Abstracts. 5th International Congress Fossils X3, Bejing, China, in press.
- A28. Saupe, E.E., Selden, P.A. & Penney, D. (2009). The first fossil Molinaranea (Araneae), with a focus on the paleobiogeography of the genus and implications for the colonization of Hispaniola. Geol. Soc. America Annual Meeting, USA, Paper No. 73-11. Abstract online at: http://gsa.confex.com/gsa/2009AM/finalprogram/abstract_164662.htm
- A27. Arnedo, M., Benjamin, S.P., Bond, J. Coddington, J., Goloboff, P., Grismado, C., Griswold, C., Hayashi, C., Hedin, M., Hormiga, G., Maddison, W., Miller, J., Platnick, N., Penney, D., Prendini, L., Ramírez, M., Scharff, N., Silva, D., Sierwald, P., Vink , C. & Wheeler, W. (2007). Spinning a new web in spider systematics: progress towards the phylogeny of Araneae. Abstracts. XVII Int. Cong. Arachnol., Brazil, 66.
- A26. Dunlop, J.A., Penney, D., Tetlie, O.E. & Anderson, L.I. (2007) How many species of fossil arachnid are there? Abstracts. XVII Int. Cong. Arachnol., Brazil, 36. Invited plenary lecture
- A25. Penney, D. (2007) Continental affinities of Hispaniolan spiders provide support for GAARlandia landspan. Abstracts. NESCent (NSF) Synthesis Meeting on West Indian Biodiversity, Dominican Republic, July 2007. Invited speaker
- A24. Penney, D. & Langan, A.M.L. (2006) Comparing amber fossil assemblages across the Cenozoic. Abstracts. 23rd European Colloq. Arachnol., Spain, 2006, 89.
- A23. Dunlop, J.A., Harms, D. & Penney, D. (2006) A fossil tarantula (Araneae: Theraphosidae) from Tertiary Chiapas amber, Mexico. Abstracts. 23rd European Colloq. Arachnol., Spain, 2006, 36.
- A22. Penney, D. & Langan, A.M.L. (2006) Comparing amber fossil assemblages across the Cenozoic. Abstracts. 9th Int. Symp. Mes. Terr. Ecosyst., UK, insert.
- A21. Jepson, J.E. & Penney, D. (2006) Neuropteran (Insecta) palaeodiversity with predictions for the Cretaceous fauna of the Wealden. Abstracts. 9th Int. Symp. Mes. Terr. Ecosyst., UK, insert.
- A20. Penney, D. (2005) Completeness and bias versus adequacy of the spider fossil record. Abstracts. 3rd Int. Palaeoentomol. Soc. Cong., South Africa, 2005, 26. Invited keynote
- A19. Penney, D. (2005) Hispaniolan spiders: extinct and extant compared. Abstracts. 3rd Int. Palaeoentomol. Soc. Cong., South Africa, 2005, 49.
- A18. Menon, F., Penney, D., Selden, P.A. & Martill, D.M. (2005) A scolopendromorph centipede from the Cretaceous Crato Formation of Brazil. Abstracts. 3rd Int. Palaeoentomol. Soc. Cong., South Africa, 2005, 48.
- A17. Penney, D. & Selden, P.A. (2005) Assembling the tree of life—phylogeny of spiders: a review of the strictly fossil spider families. Abstracts. 22nd European Colloq. Arachnol., Bulgaria, 2005, 37.
- A16. Penney, D. (2005) Hispaniolan spiders: extinct and extant compared. Abstracts. 8th African Arachnol. Colloq., South Africa, 2005, 39.
- A15. Penney, D. & Selden, P.A. (2005) Recent advances in palaeoarachnology and the true identity of the giant fossil ‘spider’ Megarachne. Abstracts. 8th African Arachnol. Colloq., South Africa, 2005, 34.
- A14. Penney, D. (2004) The enigmatic fossil spider family Lagonomegopidae in Cretaceous ambers with descriptions of new specimens from Myanmar. Abstracts. XVI Int. Cong. Arachnol., Belgium, 125.
- A13. Penney, D. (2004) Hispaniolan spiders: extinct and extant compared. Abstracts. XVI Int. Cong. Arachnol., Belgium, 260.
- A12. Penney, D.(2003) Estimations of global extant spider family richness using neontological and palaeontological data. Abstracts 21st European Colloquium of Arachnology, St. Petersburg, Russia, 67.
- A11. Penney, D., Wheater, C.P. & Selden, P.A. (2003) Resistance of spiders to Cretaceous–Tertiary extinction events. Abstracts Palass 47th Ann. Meet., Leicester, UK, 148.
- A10. Marusik, Y.M. & Penney, D. (2003) A survey of Baltic amber Theridiidae (Araneae) inclusions with descriptions of six new species. Abstracts 27th Ann. Meet. American Arachnol. Soc., Denver, Colorado. Online at http://www.americanarachnology.org/AAS_Meetings/AAS_03_abstracts.html
- A9. Marusik, Y.M. &Penney, D. (2003) A survey of Baltic amber Theridiidae (Araneae) inclusions with descriptions of six new species. Abstracts 21st European Colloquium of Arachnology, St. Petersburg, Russia, 57.
- A8. Menon, F., Penney, D., Selden, P.A. & Martill, D.M. (2003) A scolopendromorph centipede from the Cretaceous Crato Formation of Brazil. Abstracts Palass 47th Ann. Meet., Leicester, UK, 180–181.
- A7. Penney, D. (2002) Spider–insect (predator–prey) co-radiation—what does the fossil record tell us?In Abstracts: Spiders and Insects: a joint meeting of the British Arachnological Society and Royal Entomological Society, 3rd April 2002. Newsl. Br. arachnol. Soc. 95, 4–5.
- A6. Selden, P.A. & Penney, D. (2002) Mesozoic Araneae: exploring the dark ages of spider evolution. Abstracts. 8th Int. Symp. Mes. Terr. Ecosyst., South Africa, 44.
- A5. Penney, D. (2001) Spinning with the dinosaurs—Spiders in Upper Cretaceous amber from New Jersey (Arthropoda, Araneae). Abstracts. XV Int. Cong. Arachnol., South Africa, 127.
- A4. Selden, P.A. & Penney, D. (2001) Phylogeny of Araneae: the fossil evidence and its interpretation. Abstracts. XV Int. Cong. Arachnol., South Africa, 157.
- A3. Selden, P.A. & Penney, D. (2001) Phylogeny of Araneae: the fossil evidence and its interpretation. Abstracts. Palass. 45th Ann. Meet., Copenhagen, 48.
- A2. Penney, D. (1998) Hypotheses for the Recent Hispaniolan spider fauna based on the Dominican Republic amber spider fauna. Abstracts. XIV Int. Cong. Arachnol., Chicago, 28.
- A1. Penney, D. (1995) Oecobius amboseli (Oecobiidae): Description of the male, redescription of the female, and observations of intraspecific variation of guanine distribution. Abstracts. XIII Int. Cong. Arachnol., Geneva, 42.
7. Book reviews
- BR6. Penney, D. (in prep.) Review of: Hannah, L. (ed.) 2012. Saving a Million Species: extinction risk from climate change. Island Press, London. TBA, XX: XX–XX.
- BR5. Penney, D. (in press) Review of: Gordh, G. & Headrick, D. 2011. A dictionary of entomology, second edition. CSIRO Publishing, Victoria, Australia. Antenna, XX: XX–XX.
- BR4. Penney, D. (in press) Palaeoenvironmental Habitat Reconstructions: A palaeoentomological perspective. Antenna, XX: XX–XX.
- BR3. Penney, D. (in press) Palaeoenvironmental Habitat Reconstructions: A palaeoentomological perspective. Palaeont. Assoc. Newsl., XX: XX–XX.
- BR2. Omelko, M.M. & Penney, D. (2011) A long-awaited book about spiders from the Asian part of Russia. Review of: Marusik, Yu.M., Kovblyuk, M.M. 2011. Spiders (Arachnida, Aranei) of Siberia and the Russian Far East. KMK Scientific Press Ltd, Moscow. ZooKeys, 154: 89–93.
- BR1. Penney, D. (2011) Book review: Fossil Behavior Compendium by A.J. Boucot & G.O. Poinar Jr. (2010). Palaeont. Assoc. Newsl., 78: 79–82.
8. Other publications
- O44. Penney, D. (in press) New book on fossil Neuroptera? Lacewing News, XX: XX–XX.
- O43. Penney, D. & Dunlop, J.A. (submitted) Fossil Arachnids. Deposits Magazine, XX: XX–XX. Invited paper
- O42. Dunlop, J.A. & Penney, D. (2012) Fossilspinnen. SENCKENBERG Natur Forschung Museum, 142: 120–125. Invited paper
- O41. Penney, D. (2011) Nematodes of the Seychelles Islands. Nematology Newsletter, 57: 11.
- O40. Penney, D. (2011) Out of the forest: Gambian arachnids. Micro Miscellanea, 78: 6–9.
- O39. Penney, D. & Green, D.I. (in press) Sub-fossils in copal: an undervalued resource.Deposits Magazine, XX: XX–XX, Invited paper
- O38. Penney, D. & Green, D.I. (in press) Palaeodiversity of fossil inclusions in Dominican Amber with an emphasis on Hexapoda. For Antenna. Invited paper
- O37. Penney, D. & Green, D.I. (2011) Fossil amber inclusions under the microscope. Micro Miscellanea, 77: 18–21.
- O36. Penney, D. (2011) New techniques for studying fossil spiders in amber. Newsl. Br. arachnol. Soc., 121: 1–5.
- O35. Penney, D. & Green, D.I. (2011) Preparation and study of fossils in amber.Deposits Magazine, 27: 10–16.Invited paper
- O34. Penney, D. & Green, D.I. (2011) Biodiversity of fossils in amber.Deposits Magazine, 26: 42–48. Invited paper
- O33. Penney, D. (2009) Unexpected synanthropic spiders in The Gambia, West Africa (Araneae: Cithaeronidae, Prodidomidae). Newsl. Br. arachnol. Soc., 116: 6–7.
- O32. Penney, D. & Bibi, Z. (2007) Spiders focus in synthesis of Caribbean biodiversity. Newsl. Br. arachnol. Soc. 108, 6–7.
- O31. Penney, D. (2005) Hispaniolan spiders: extinct and extant compared. Abstract.Newsl. African Arachnol. Soc.18, 25.
- O30. Penney, D.& Selden, P.A. (2005) Recent advances in palaeoarachnology and the true identity of the giant fossil ‘spider’ Megarachne. Abstract only. Newsl. African Arachnol. Soc.18, 23–24.
- O29. Penney, D. (2005) Two recent arachnological meetings in South Africa. Newsl. Br. arachnol. Soc. 103, 6–7.
- O28. Penney, D. (2005) Toads and spiders. Newsl. Br. arachnol. Soc. 102, 2.
- O27. Penney, D. (2004) A palaeoarachnological trip to Paris. Newsl. Br. arachnol. Soc. 101, 13–14.
- O26. Penney, D. (2004) A rather unusual spider book. Newsl. Br. arachnol. Soc. 99, 5–6.
- O25. Logunov, D.V. & Penney, D. (2004) A post-colloquium trip to Ciscaucasia (Russia). Newsl. Br. arachnol. Soc99, 4–5.
- O24. Penney, D. (2003) 21st European Colloquium of Arachnology, St. Petersburg, Russia, 4–9 August 2003. Newsl. Br. arachnol. Soc. 98, 14–15.
- O23. Penney, D. (2003) To the Dominican Republic in Search of Amber Spiders. Newsl. Br. arachnol. Soc. 97, 7–8.
- O22. Penney, D. (2003) A taste of Russia in Manchester. Newsl. Br. arachnol. Soc. 96, 19–20.
- O21. Penney, D. (2003) No such spider. New Sci.178(2378), 24.
- O20. Penney, D. (2002) Age of spider-inclusion-bearing Cretaceous ambers: an update. Newsl. Br. arachnol. Soc. 95, 16.
- O19. Penney, D. (2002) Arachnological gems: spiders in amber. Newsl. Br. arachnol. Soc. 94, 2–5.
- O18. Penney, D. (2001) Amazonian spiders: a reply. Newsl. Br. arachnol. Soc. 91, 7.
- O15. Penney, D. (1999) Dominican Republic amber spiders and their contribution to fossil and Recent ecology. Unpublished PhD thesis, Department of Earth Sciences, The University of Manchester, 377pp.
- O16. Penney, D. (1999) XIV International Congress of Arachnology—The Field Museum of Natural History, Chicago, Illinois, 27 June–3 July 1998. Meganeura, 3, 2–3.
- O15. Penney, D. (1998) Doctoral thesis abstract—Dominican Republic amber spiders and their contribution to fossil and Recent ecology. Meganeura, 2, 24–25.
- O14. Penney, D. (1997) Can scorpions jump to avoid predation? Newsl. Br. arachnol. Soc. 79, 3.
- O13. Penney, D. (1997) Spider quest to Panama. Newsl. Br. arachnol. Soc. 78, 4–5.
- O12. Penney, D. (1996) Micropholcus fauroti (Simon, 1887) A new spider record for Panama. Newsl. Br. arachnol. Soc., 77, 11.
- O11. Penney, D. & Drew, B. (1996) A novel technique for extracting Segestria spp. from their retreats. Newsl. Br. arachnol. Soc., 76, 2–3.
- O10. Penney, D. (1996) Notes on Hersilia occidentalis Simon from Uganda and its possible association with ants. Newsl. Br. arachnol. Soc., 75, 5–6.
- O9. Penney, D. (1995) Restrictions on research on, and collection of spiders from Uganda. Newsl. Br. arachnol. Soc., 74, 12.
- O8. Penney, D. (1995) No tuning-fork? Try forceps. Newsl. Br. arachnol. Soc., 74, 10.
- O7. Penney, D. (1995) Theridion pallens, another reply. Newsl. Br. arachnol. Soc., 74, 2–3.
- O6. Penney, D. (1995) Spider quest to Uganda. Newsl. Br. arachnol. Soc., 72, 7.
- O5. Penney, D. & Whitehead, J. (1995) Note on Deinopis schoutedeni Giltay from Rubaga, Uganda. Newsl. Br. arachnol. Soc., 73, 2–3.
- O4. Penney, D. & Ramsay, J. (1994) A Meta menardi up a tree. Newsl. Br. arachnol. Soc., 70, 7–8.
- O3. Jones-Walters, L., Dalingwater, J. & Penney, D. (1993) Notes on the spiders of Bernac and the adjacent area, France. Newsl. Br. arachnol. Soc., 68, 5–7.
- O2. Varagona, G., Ellis, L.A., Moore, D, Penney, D, & Dusheiko, G.M. (1991) A percutaneous liver biopsy technique in ducks (Anas platyrhynchos) experimentally infected with duck hepatitis B virus. Lab. Anim., 25, 254–257.
- O1. Penney, D. (1986) A note on the Orthognath spider Melopeus albostriatus. Newsl. Br. arachnol. Soc., 47, 5.
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